species had left behind it in its native 
European or African mainland. 
I said a little while ago we had no mammal in the islands. In that I was 
not quite strictly correct. I ought to have said, no terrestrial mammal. A 
little Spanish bat got blown to us once by a rough nor'easter, and took 
up its abode at once among the caves of our archipelago, where it 
hawks to this day after our flies and beetles. This seemed to me to show 
very conspicuously the advantage which winged animals have in the 
matter of cosmopolitan dispersion; for while it was quite impossible for 
rats, mice, or squirrels to cross the intervening belt of three hundred
leagues of sea, their little winged relation, the flitter-mouse, made the 
journey across quite safely on his own leathery vans, and with no 
greater difficulty than a swallow or a wood-pigeon. 
The insects of my archipelago tell very much the same story as the 
birds and the plants. Here, too, winged species have stood at a great 
advantage. To be sure, the earliest butterflies and bees that arrived in 
the fern-clad period were starved for want of honey; but as soon as the 
valleys began to be thickly tangled with composites, harebells, and 
sweet-scented myrtle bushes, these nectar-eating insects established 
themselves successfully, and kept their breed true by occasional crosses 
with fresh arrivals blown to sea afterwards. The development of the 
beetles I watched with far greater interest, as they assumed fresh forms 
much more rapidly under their new conditions of restricted food and 
limited enemies. Many kinds I observed which came originally from 
Europe, sometimes in the larval state, sometimes in the egg, and 
sometimes flying as full-grown insects before the blast of the angry 
tempest. Several of these changed their features rapidly after their 
arrival in the islands, producing at first divergent varieties, and finally, 
by dint of selection, acting in various ways, through climate, food, or 
enemies, on these nascent forms, evolving into stable and well-adapted 
species. But I noticed three cases where bits of driftwood thrown up 
from South America on the western coasts contained the eggs or larvae 
of American beetles, while several others were driven ashore from the 
Canaries or Madeira; and in one instance even a small insect, belonging 
to a type now confined to Madagascar, found its way safely by sea to 
this remote spot, where, being a female with eggs, it succeeded in 
establishing a flourishing colony. I believe, however, that at the time of 
its arrival it still existed on the African continent, but becoming extinct 
there under stress of competition with higher forms, it now survives 
only in these two widely separated insular areas. 
It was an endless amusement to me during those long centuries, while I 
devoted myself entirely to the task of watching my fauna and flora 
develop itself, to look out from day to day for any chance arrival by 
wind or waves, and to follow the course of its subsequent vicissitudes 
and evolution. In a great many cases, especially at first, the new-comer
found no niche ready for it in the established order of things on the 
islands, and was fain at last, after a hard struggle, to retire for ever from 
the unequal contest. But often enough, too, he made a gallant fight for 
it, and, adapting himself rapidly to his new environment, changed his 
form and habits with surprising facility. For natural selection, I found, 
is a hard schoolmaster. If you happen to fit your place in the world, you 
live and thrive, but if you don't happen to fit it, to the wall with you 
without quarter. Thus sometimes I would see a small canary beetle 
quickly take to new food and new modes of life on my islands under 
my very eyes, so that in a century or so I judged him myself worthy of 
the distinction of a separate species; while in another case, I remember, 
a south European weevil evolved before long into something so wholly 
different from his former self that a systematic entomologist would 
have been forced to enrol him in a distinct genus. I often wish now that 
I had kept a regular collection of all the intermediate forms, to present 
as an illustrative series to one of your human museums; but in those 
days, of course, we none of us imagined anybody but ourselves would 
ever take an interest in these problems of the development of life, and 
we let the chance slide till it was too late to recover it. 
Naturally, during all these ages changes of other sorts were going on in 
my islands--elevations and subsidences, separations and reunions, 
which helped to modify the life of the group considerably. Indeed, 
volcanic action was constantly at work altering the    
    
		
	
	
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